They find out about each other, and realize that they each have a 50/50 chance of being the twin. Each girl came believing that she was the twin sister. In fact, in the Arabian Nights, a man will call a woman he loves or is married to "daughter of my uncle", because in that time marriage between cousins was considered the best option! Because of this, it almost never appears as a plot point since it's a little too close for some folks to be comfortable, but not close enough that the plot demands the relationship break up. Cousins are only borderline squicky since marriage between cousins is actually legal in some regions, and is quite acceptable in many romance novels. The surprise relationship is usually either brother/sister, half-siblings, parent/child (much rarer these days than it was in ancient Greece), or, very rarely, uncle/niece or aunt/nephew. Still, this trope is surprisingly common in Anime and Manga, as well as other Japanese entertainment, due to much more relaxed views on the subject in Japan.
This wasn't always the case Oedipus was very happily (but unknowingly) married to his own mother for many years and had several kids with her before things fell apart (to the extent that The Reveal resulted in Oedipus's mother killing herself and Oedipus himself stabbing his own eyes out), so this is Older Than Feudalism. These days, many people are squicked out by this, so the characters often find out before they tie the knot/do the nasty. Or if the two still want to be together and the author doesn't want to go the incest route, a counter-reveal may show up to prove that the genetics aren't a problem after all. If it's not called off despite the consanguinity issue, intentional Brother-Sister Incest results. The result may be either great rejoicing or great despair. But then, someone makes a startling discovery - genetics has struck! The couple-to-be are related - and not in the "same species" sense - and the romance or arranged marriage is called off.
Love blossoms in a particular way, or perhaps the two have a loveless Arranged Marriage lying in wait for them instead. Sometimes in stories, two people meet and just seem destined for romance. The Game The Whole Family Can Play! Haha! Guess what you were doing?
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